Turbellarian taxonomic database

Acoela Haploposthiidae


Westblad E 1948 (citation)- p 52 sets up this family as having "a ventral mouth without a pharynx
	combined with Proporidae characteristic reproductive system." 

	p 55-56, key as follows:

	Tribe Opisthandropora- Abursalia
	Family Haploposthidae- "Mouth ventrl.  Pharynx open.  Male and female gonads with respect to
		position and appearance variable but never combined into a compact gondad unit.
		
		A1- Male antrum +- long and tube shaped.  
			B1- Reproductive glands [gonads] often mixed but the 
				female +- lying dorsally----------------------------------genus Haploposthia
				
			B2- male and female gondads separate, both the male and the female paired;
				the female gonad lying ventrally--------------------------genus Paranaperus
				
		A2- Male antrum short
			B1- A short penis; small and +- round in cross section
				C1- Frontal glands small (not "a frontal organ"); ovary unpaired;
					copulatory organ with weak cutecularized needles------genus Paraproporus
					
				C2- Frontal glands lacking.  Ovaries paired.  Copulatory organs without
					cuticular needles-------------------------------------genus Afronta (Hyman)
					
			B2- Double, stylet formed penis.  Fairly large, flat shape----genus Childia
			
Marcus E 1950 (citation)- points out family is characterized by absense of a pharynx.  genera belonging
	to it are characterized as follows:
		Haploposthia- colored rhabdite gland
		Paranaperus- a penis and spacious atrium
		Paraproporus- by penis rods
		Childia- double copulatory organs
		Afronta- 1 species only with orange spots
		Kuma- large frontal glands, short atrium and no penis cuticularized atrium.
										
Dorjes J 1968 (citation)- p 94-95.  Definition- "Acoels with very simple copulatory organs.  These 
	consist of a tube-shaped male antrum and a keg-like seminal vesicle.  Both the antrum as
	well as the vesicle can be mutually or entirely lacking.  As penis is not formed 
	[Haploposthia albiventer Marcus and H. rubra (An der Lan) with an extremely small penis
	papilla].  Male reproductive opening terminal, subterminal or ventral.  Mouth opening ventral."
	Type of family: genus Haploposthia An der Lan 1936- 11 genera with 32 species.
	
	1.  Haploposthia		7.  Parahaplogonaria
	2.  Afronta				8.  Parahaploposthia
	3.  Kuma				9.  Pseudohaplogonaria
	4.  Adenocauda			10. Pseudokuma
	5.  Deuterogonaria 		11. Simplicomorpha
	6.  Haplogonaria		
	
	Discussion of family on p 95, p 95-96 key to genera.
	
Faubel A 1976 (citation)- lists.
Ehlers U, Doerjes J 1979 (citation)- discussion p 29, 48, and 70.
Faubel A 1983 (citation)- p 26, notes ciliated male tract here.


Raikova OI 1989 (citation)- "The fine structure of the nervous system of both plexus-type and that with formed
nerve trunks was studied in representatives of four families of Acoela."  Families and species studied
include:
     Haploposthiidae
        Haploposthia opisthorchis
     Childiidae
        Actinoposthia beklemischevi
     Anaperidae
        Anaperus biaculeatus
     Convolutidae
        Aphanostoma virescens

Descriptions of synapses, vesciles with granules in neurites, sensillae as associated fine structure are
described.