tetraculeatus

	site	site # (info)	collection date	kind	depth	substrate	salin	comments	reference
A	Nielönn, north of Kampen, Sylt Island, Germany, North Sea	3613 map	1982-1983	type locality				salt meadow. 26 individuals.	Armonies W (1987): 98
B	Nielönn, north of Kampen, Sylt Island, Germany, North Sea	3613 map	Dec 1982 - Jul 1983	type locality				Mittlerer Andelrasen der Sazwiese "Nielönn", 26 Individuen.	Armonies W, Hellwig M (1987): 252