|  | site | site # (info) | collection date | kind | depth | substrate | salin | comments | reference | 
| A | Lake Glensevern and Upper Duck Pond, Campobello Island, New Brunswick, Canada | 3589 
  map | Aug-Sep 1984 |  |  | medium-coarse sand |  | Upper Duck Pond.  Intertidal medium to coarse sand with halophytes. | Ax P, Armonies W (1987): 20 | 
| B | Weser River Estuary, Lower Saxony, Germany | 3656 
  map | 1993-1994 |  |  | medium-coarse sand |  | Weser Estuary (Germany, Lower Saxony) samples taken from intertidal flats.  Mittel- und Grobsand im Meso- und Polyhalinicum. | Krumwiede A, Witt J (1995): 321 | 
| C | Weddewarden, Weser River Estuary, Lower Saxony, Germany | 3663 
  map | 1993-1994 |  |  |  |  | Burhave (Nordbereich des Strandbades Burhave), Weser Estuary samples taken from intertidal flats. | Krumwiede A, Witt J (1995): 322 | 
| D | Sahlenburg Watt, Weser River Estuary, Lower Saxony, Germany | 3665 
  map | 1993-1994 |  |  |  |  | Sahlenburg (Hundestrand), Weser Estuary samples taken from intertidal flats. | Krumwiede A, Witt J (1995): 322 | 
| E | De Panne (La Panne), Belgium | 4175 
  map | Aug 1981 | type locality |  | fine sand |  | Belgium North-Sea coast, De Panne, mediolittoral. | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 252 | 
| F | De Panne (La Panne), Belgium | 4175 
  map | May 1983 | type locality |  | fine sand |  | Belgium North-Sea coast, De Panne, mediolittoral. | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238, 252 | 
| G | Oostende Bank, Belgium | 3543 
  map | May 1983 |  |  |  |  |  | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238 | 
| H | Het Zwin (Zwin Estuary, Zwijn), Belgium | 4176 
  map | May 1983 |  |  |  |  | Zwin (beach). | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238 | 
| I | Het Zwin (Zwin Estuary, Zwijn), Belgium | 4176 
  map | Jan 1984 |  |  |  |  | Zwin (beach).  Collector Revis. | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238 | 
| J | Het Zwin (Zwin Estuary, Zwijn), Belgium | 4176 
  map | Mar 1984 |  |  |  |  | Zwin (beach).  Collector Revis. | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238 | 
| K | Heist (Heyst, Heist-sur-Mer, Heist-aan-Zee), Belgium | 3947 
  map | Oct 1983 - Feb 1984 |  |  |  |  | Heist.  Collector Jouk. | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238 | 
| L | Sandy beach on eastern shore of Sylt.  Litoralstation (old littoral station), Biological Institute Helgoland, Biologische Anstalt Helgoland (BAH), List, Sylt | 323 
  map | Sep 1985 |  |  |  |  | medium sand in the mediolitoral of beaches along the Belgium coast, near the littoral station of Sylt. | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238, 252 | 
| M | Marine Biological Research Station Roscoff, Roscoff (Roskoff),  France | 3315 
  map | Jul 1986 |  |  |  |  | in front of the Biological Station. | Martens PM, Curini-Galletti MC, Puccinelli I (1989): 238, 252 | 
| N | Belgium coast | 5471 
  map | 1989 or earlier |  |  |  | brackish | Die Art dringt in Brackgewässer vor:  Belgien. | Schockaert ER, Jouk PEH, Martens PM (1989): ; from Ax 2008 (citation)- p. 219 |