History of
  Turbellarian Research
      compiled by Steve Schilling
Listing of publications with author Sluys R
author date title journal / publication
Agapow PM, Sluys R 2005 The reality of taxonomic change. Trends in Ecology & Evolution 20: 278-280.
Aliabadian M, Roselaar CS, Nijman V, Sluys R, Vences M 2005 Identifying contact zone hotspots of passerine birds in the Palaearctic region. Biology Letters 1: 21-23.
[doi: doi.org/10.1098/rsbl.2004.0258
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Aliabadian M, Roselaar CS, Nijman V, Sluys R, Vences M 2006 The mid-domain effect on the pattern of geographic species richness in Palearctic songbirds. Journal of Ornithology 147: 80-80.
Aliabadian M, Roselaar CS, Sluys R, Nijman V 2007 Low predictive power of mid-domain effect to explain geographic species richness patterns in Palearctic songbirds. Contributions to Zoology 76: 197-204.
Aliabadian M, Sluys R, Roselaar CS, Nijman V 2008 Species diversity and endemism: testing the mid-domain effect on species richness patterns of songbirds in the Palearctic Region. Contributions to Zoology 77: 99-108.
Álvarez-Presas M, Mateos E, Sluys R, Riutort M 2022 Cryptic diversity in European terrestrial flatworms of the genus Microplana (Platyhelminthes, Tricladida, Geoplanidae) Cryptic Species : 281-293
[doi: 10.1017/9781009070553.011
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Alvarez-Presas M, Mateos E, Vila-Farre M, Sluys R, Riutort M 2012 Evidence for the persistence of the land planarian species Microplana terrestris (Muller, 1774) (Platyhelminthes, Tricladida) in microrefugia during the Last Glacial Maximum in the northern section of the Iberian Peninsula. Molecular Phylogenetics and Evolution 64: 491-499.
[doi: doi.org/10.1016/j.ympev.2012.05.001
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Benitez-Alvarez L, Leal-Zanchet AM, Oceguera-Figueroa A, Ferreira RL, Bento D de M, Braccini J, Sluys R, Riutort M 2020 Phylogeny and biogeography of the Cavernicola (Platyhelminthes: Tricladida): Relicts of an epigean group sheltering in caves? Molecular Phylogenetics and Evolution, Volume: 145: 106709
Benitez-Alvarez L, Leria L, Sluys R, Leal-Zanchet AM, Riutort M 2023 Niche modelling and molecular phylogenetics unravel the colonisation biology of three species of the freshwater planarian genus Girardia (Platyhelminthes, Tricladida). Hydrobiologia
[https://doi.org/10.1007/s10750-023-05239-x
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Benitez-Alvarez L, Sluys R, Leal-Zanchet AM, Leria L, Riutort M 2023 First molecular phylogeny of the freshwater planarian genus Girardia (Platyhelminthes: Tricladida) unveils hidden taxonomic diversity and initiates resolution of its historical biogeography. Zoological Journal of the Linnean Society, Vol 197 (1): 1-19
[doi: 10.1093/zoolinnean/zlac065
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Caceres-Martinez J, Vasquez-Yeomans R, Sluys R 1996 Urastoma cyprinae in natural and cultured Mussel (Mytilus galloprovincialis Lmk) populations in México. Bulletin of the European Association of Fish Pathologists 16: 200-202.
Caceres-Martinez J, Vasquez-Yeomans R, Sluys R 1998 The turbellarian Urastoma cyprinae from edible mussels Mytilus galloprovincialis and Mytilus californianus in Baja California, NW Mexico. J Invert Pathol 72: 214-219
Charni M, Harrath AH, Sluys R, Tekaya S, Zghal F 2004 The freshwater planarian Dugesia sicula Lepori, 1948 (Platyhelminthes, Tricladida) in Tunisia: ecology, karyology, and morphology. Hydrobiologia 517: 161-170
Chen G, Wang L, Wu F, Sun X, Dong Z, Sluys R, Yu F, Yu-wen Y, Liu D 2022 Two new species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from the subtropical monsoon region in Southern China, with a discussion on reproductive modalities BMC Zoology 7(1):
[doi: 10.1186/s40850-022-00127-8
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Chen JJ, Li W-X, Sluys R, Wu M-Q, Wang L, Li S-F, Wang A-T 2019 Two new species of marine flatworm from southern China facilitate determination of the phylogenetic position of the genus Nerpa Marcus, 1948 and the histochemical structure of the nervous system in the genus Paucumara Sluys, 1989 (Platyhelminthes, Tricla Zootaxa, Volume:4568(1): 149-167
De Vries EJ, Sluys R 1991 Phylogenetic relationships of the genus Dugesia (Platyhelminthes, Tricladida, Paludicola). J Zool Lond 223: 103-116. (Abstr kl Karte)
De Waart SA, Thunnissen NW, Sluys R 2021 Nieuwe vondsten van exotische landplatwormen, waaronder vijf nieuwe soorten voor Nederland (Platyhelminthes: Geoplanidae). [New records of exotic land flatworms, with five new species for the Netherlands (Platyhelminthes: Geoplanidae).] Nederlandse Faunistische Mededelingen 56: 15–28 [in Dutch, with English summary]
Dols-Serrate D, Stocchino GA, Sluys R, Riutort M 2023 Fantastic beasts and how to delimit them: an integrative approach using multispecies coalescent methods reveals two new, endemic Dugesia species (Platyhelminthes: Tricladida) from Corsica and Sardinia Zoological Journal of the Linnean Society :
[doi: 10.1093/zoolinnean/zlad135
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Dorigo L, Lago T D, Menchetti M, Sluys R 2020 First records of two alien land flatworms (Tricladida, Geoplanidae) from Northeastern Italy. Zootaxa 4732: 332–334
[https://doi.org/10.11646/zootaxa.4732.2.8
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Estep K, Sluys R, Syvertsen EE 1993 "Linnaeus" and beyond: workshop report on multimedia tools for the identification and database storage of biodiversity. Hydrobiologica 269/270: 519-525
Faubel A, Sluys R, Reid DG 2007 A new genus and species of polyclad flatworm found in the mantle cavities of gastropod molluscs in the high-intertidal zone of the Pacific coast of central America. Journal of the Marine Biological Association of the United Kingdom 87(2): 429-434.
Gibbons MJ, Sluys R 1999 The taxonomic richness of South Africa's marine fauna: a crisis at hand. South African Journal of Science 95: 8-12.
Grant LJ, Sluys R, Blair D 2006 Biodiversity of Australian freshwater planarians (Platyhelminthes: Tricladida: Paludicola): new species and localities, and a review of paludicolan distribution in Australia. Systematics and Biodiversity 4(4): 435-471
Grau JH, Almeida AL, Sluys R, Carbayo F 2022 A new genus and two new species of land planarians (Platyhelminthes: Tricladida: Geoplanidae) from Southern Chile. Journal of Natural History, Volume 56, 2022 - Issue: 13-16
[https://doi.org/10.1080/00222933.2022.2097137
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Grau JH, Sluys R, Froehlich EM, Carbayo F 2012 Reflections on the genus Amaga Ogren and Kawakatsu 1990, and description of a new genus of land planarian (Platyhelminthes: Tricladida: Geoplanidae) Journal of Natural History 46: 1529-1546
[doi: 10.1080/00222933.2012.691996
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Harrath AH, Sluys R, Mansour L, Folefack GL, Aldahmash W, Alwasel S, Sola E 2019 Molecular and morphological identification of two new African species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from Cameroon. Journal of Natural History, Volume:53(5-6): 253-271
Harrath AH, Charni M, Sluys R, Zghal F, Tekaya S 2004 Ecology and distribution of the freshwater planarian Schmidtea mediterranea in Tunisia. Ital J Zool 71:233-236
Harrath AH, Mansour L, Lagnika M, Sluys R, Boutin C, Alwasel S, Poch A, Riutort M 2016 A molecular analysis of the phylogenetic position of the suborder Cavernicola within the Tricladida (Platyhelminthes), with the description of a new species of stygobiont flatworm from Benin Zoological Journal of the Linnean Society, Volume 178, Issue 3, November 2016, Pages 482–491
[doi: doi.org/10.1111/zoj.12430
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Harrath AH, Sluys R, Aldahmash W, Al-Razaki A, Alwasel S 2013 Reproductive Strategies, Karyology, Parasites, and Taxonomic Status of Dugesia Populations from Yemen (Platyhelminthes: Tricladida: Dugesiidae). Zoological Science (Tokyo), 30(6): 502-508
Harrath AH, Sluys R, Ghlala A, Alwasel S 2012 The first subterranean freshwater planarians from North Africa, with an analysis of adenodactyl structure in the genus Dendrocoelum (Platyhelminthes, Tricladida, Dendrocoelidae) Journal of Cave and Karst Studies 74(1): 48–57.
[doi: 10.4311/2011LSC0215
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Harrath AH, Sluys R, Merzoug D, Yacoubi-Khebiza M, Alwasel S, Riutort M 2012 Freshwater planarians (Platyhelminthes, Tricladida) from the Palearctic section of the African continent: new records, with the description of a new species ZOOTAXA
Harrath AH, Sluys R, Zghal F, Tekaya S 2009 First report of adelphophagy in flatworms during the embryonic development of the planarian Schmidtea mediterranea (Benazzi, Baguna, Ballester, Puccinelli & Del Papa, 1975) (Platyhelminthes, Tricladida). Invertebrate Reproduction and Development 53: 117-124.
[doi: doi.org/10.1080/07924259.2009.9652297
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Hovenkamp P, Gittenberger E, Hennipman E, Jong R de, Roos MC, Sluys R, Zandee M 1987 Systematics and Evolution: A Matter of Diversity. (editors) Utrecht University, ISBN 9789090019673.; 341 pages
Kawakatsu M, Mitchell RW, Sluys R, Sasaki GY 2002 A list of triclad turbellarians from Mexico in Kawakatsu's collection. Kawakatsu and Sasaki's Webpages on Planarians. 2002 Aug 17; 37:1-10. http://planarian.net/kswp/37/mexico.pdf
Kawakatsu M, Sluys R, Ogren R 2005 Seven new species of land planarian from Japan and China (Platyhelminthes, Tricladida, Bipaliidae), with a morphological review of all Japanese bipaliids and a biogeographic overview of Far Eastern species. Belgian Journal of Zoology 135(1): 53-77
Kawakatsu M, Sluys R, Sasaki GY 2004 Type series of Japanese freshwater "Turbellarians". Kawakatsu & Sasaki's WEBPAGES ON PLANARIANS, SAPPORO AND TÔKYÔ, No. 44, Apr 13, 2004. pp 1-25 [http://planarian.net/]
Kawakatsu M, Sluys R, Timoshkin OA, Naumova TV, Nishino M, Takai M 2000 Redescription of Bdellocephala annandalei, an endemic dendrocoelid planarian from Lake Biwa-ko in Honshu, Central Japan. (Turbellaria, Tricladida, Paludicola). Preprint 9th ISBT
Kawakatsu M, Sluys R, Timoshkin OA, Naumova TV, Nishino M, Takai M 2001 Redescription of Japanese Bdellocephala annandalei from Lake Biwa-ko with comparative redescription of the Far Eastern and Kamchatkan Bdellocephala species (Tricladida, Paludicola). Belg J Zool 131 (Suppl 1): 205-211
Lazaro EM, Sluys R, Pala M, Stocchino GA, Baguñà J, Riutort M 2009 Molecular barcoding and phylogeography of sexual and asexual freshwater planarians of the genus Dugesia in the Western Mediterranean (Platyhelminthes, Tricladida, Dugesiidae) Molecular Phylogenetics and Evolution 52: 835-845
[doi: doi.org/10.1016/j.ympev.2009.04.022
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Leria L, Sluys R, Riutort M 2018 Diversification and biogeographic history of the Western Palearctic freshwater flatworm genus Schmidtea (Tricladida: Dugesiidae), with a redescription of Schmidtea nova Journal of Zoological Systematics, Vol 56 (3): 335-351
Leria L, Vila-Farré M, Álvarez-Presasa M, Sánchez-Gracia A, Roza J, Sluys R, Riutor M 2020 Cryptic species delineation in freshwater planarians of the genus Dugesia (Platyhelminthes, Tricladida): Extreme intraindividual genetic diversity, morphological stasis, and karyological variability Molecular Phylogenetics and Evolution Volume 143, February 2020, 106496; https://doi.org/10.1016/j.ympev.2019.05.010
[doi: doi.org/10.1016/j.ympev.2019.05.010
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Li M-Y, Ma X-Y, Li X-Y, Yang Y, Sluys R, Chen J-J, Li S-F, Wang A-T 2021 A new species of Pacifides from the Western Pacific Coast and the first fully freshwater species of the maricolan planarian genus Paucumara (Platyhelminthes, Tricladida, Maricola). Systematics and Biodiversity, Volume 19(5): 488-506
[https://doi.org/10.1080/14772000.2021.1877846
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Liu Y, Song X-Y, Sun Z-Y, Li W-X, Sluys R, Li -F, Wang A-T 2022 Addition to the known diversity of Chinese freshwater planarians: integrative description of a new species of Dugesia Girard, 1850 (Platyhelminthes, Tricladida, Dugesiidae). Zoosystematics and Evolution, Vol 98(2): 233-243
[doi: doi: 10.3897/zse.98.83184
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Mateos E, Sluys R, Riutort M, Álvarez-Presas M 2017 Species richness in the genus Microplana (Platyhelminthes, Tricladida, Microplaninae) in Europe: as yet no asymptote in sight. Invertebrate Systematics, 31, 269–301.
[https://doi.org/10.1071/IS16038
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Mazza G, Menchetti M, Sluys R, Sola E, Riutort M, Tricarico E, Justine J-L, Cavigioli L, Mori E 2016 First report of the land planarian Diversibipalium multilineatum (Makino & Shirasawa, 1983) (Platyhelminthes, Tricladida, Continenticola) in Europe. Zootaxa, 4067(5): 577-580
Ogren RE, Sluys R 1998 Selected characters of the copulatory organs in the land planarian family Bipaliidae and their taxonomic significance (Tricladida: Terricola). Hydrobiologia 383: 77-82
Ogren RE, Sluys R 2001 The genus Humbertium gen. nov., a new taxon of the land planarian family Bipaliidae (Tricladida, Terricola). Belg J Zool 131(Suppl 1):201-204
Robledo JAF, Caceres-Martinez J, Sluys R, Figueras A 1994 The parasitic turbellarian Urastoma cyprinae (Platyhelminthes: Urastomidae) from blue mussel Mytilus galloprovincialis in Spain: occurence and pathology. Diseases Aquatic Organisms 18:203-210
Rohde K, Watson NA, Sluys R 1990 Ultrastructure of the flame bulbs and protonephridial capillaries of Romankenkius sp. (Platyhelminthes, Tricladida, Dugesiidae). J Submicrosc Cytol Pathol 22(4): 489-496.
Roselaar CS, Sluys R, Aliabadian M, Mekenkamp PGM 2007 Geographic patterns in the distribution of Palearctic songbirds. Journal of Ornithology 148: 271-280.
Sangster G, Hazevoet CJ, Berg AB van den, Roselaar CSK, Sluys R 1999 Dutch avifaunal list: Species concepts, taxonomic instability, and taxonomic changes in 1977-1998. Ardea 87: 139-166.
Schockaert ER, Hooge M, Sluys R, Schilling S, Tyler S, Artois T 2008 Global diversity of free living flatworms (Platyhelminthes, "Turbellaria") in freshwater. Hydrobiologia 595:41-48
Slay ME, Elliott WR, Sluys R 2006 Cavernicolous Missouri triclad (Platyhelminthes: Turbellaria) records. Southwestern Naturalist 51(2): 251-252
Sluys R 1979 Notes on the characters and synonymy of Physocypria kliei Schäfer, 1934 and Potamocypris unicaudata Schäfer, 1943 (Crustacea, Ostracoda). Bulletin Zoölogisch Museum, Universiteit van Amsterdam 6: 169-176.
Sluys R 1981 Ostracods (Ostracoda, Crustacea) from two freshwater lakes near Maarssen, The Netherlands. Hydrobiological Bulletin: Netherlands Journal of Aquatic Ecology 15: 175-182.
Sluys R 1982 Bearded Tits Panurus biarmicus in The Netherlands and England: comments on the hybridization theory. Journal of Ornithology 123: 175-182.
Sluys R 1982 Geographical variation of the Kittiwake Rissa tridactyla. Gerfaut: Revue Belge d'Ornithologie 72: 221-230.
Sluys R 1983 A new species of land planarian from Madeira Portugal (Platyhelminthes, Turbellaria, Tricladida) J Zool (Lond) 201:433-444
Sluys R 1983 A new marine triclad ectoparasitic on Malaysian and Indonesian horseshoe crabs (Platyhelminthes, Turbellaria, Tricladida). Bijdr. tot de Dierkunde 53:218-226
Sluys R 1983 On a functional-morphological approach to phylogenetic reconstruction: a critique. Acta Biotheor. 32: 29-41
Sluys R 1983 Evidence for hybrid murre reconsidered. Auk 100: 236-237.
Sluys R 1983 Geographical variation and distribution of the bearded tit Panurus-biarmicus (Linnaeus, 1758) (Aves). Bijdragen tot de Dierkunde 53: 13-32.
Sluys R 1984 The meaning and implications of genealogical tree diagrams. Zeitschrift für Zoologische Systematik und Evolutionsforschung 22: 1-8.
Sluys R 1985 Various chapters (The Birds of the Western Palearctic, vol. IV). The Birds of the Western Palearctic, vol. IV: 182-183; 194-195; 206-207; 217-219; 228-229; 230; 242-243:
Sluys R 1986 Taxonomic studies on marine triclads (Turbellaria, Tricladida, Maricola). Hydrobiologia 132: 257-262
Sluys R 1986 First representative of the order Macrostomida in Australia (Platyhelminthes, Macrostomidae). Rec S Aust Mus 19(18): 399-404.
Sluys R 1987 The taxonomy of three species of the genus Procerodes (Platyhelminthes: Tricladida: Maricola). Journal of the Marine Biological Association of the United Kingdom 67: 373-384
Sluys R 1988 Structural and functional morphology within the genus Synsiphonium (Tricladida, Maricola). In: P Ax, U Ehlers, B Sopott-Ehlers (eds) Free-living and symbiotic Plathelminthes. Gustav Fischer Verlag, St Fortschritte Zool. 36:413-418
Sluys R 1988 On adaptation, the assessment of adaptations, and the value of adaptive arguments in phylogenetic reconstruction. Zeitschrift für Zoologische Systematik und Evolutionsforschung 26: 12-26.
Sluys R 1989 A monograph of marine triclads. A.A. Balkema, Rotterdam, Brookfield. 463 pp
Sluys R 1989 Phylogenetic relationships of the triclads (Platyhelminthes, Seriata, Tricladida). Bijdragen tot de Dierkunde 59:3-25
Sluys R 1989 Sperm resorption in triclads (Platyhelminthes, Tricladida). Invertebr Reprod & Develop 15(2): 89-96.
Sluys R 1989 A monograph of the marine triclads (Platyhelminthes, Seriata, Tricladida). A.A. Balkema, Rotterdam-Brookfield xii + 463 pp.
Sluys R 1989 Rampant parallelism: an appraisal of the use of nonuniversal derived character states in phylogenetic reconstruction. Syst Zool 38:350-370
Sluys R 1990 On Bdellasimilis barwicki (Platyhelminthes: Tricladida) and its phyletic position. Invertebr Taxon 4:149-158
Sluys R 1990 A monograph of the Dimarcusidae (Platyhelminthes, Seriata, Tricladida). Zool Scr 19:13-29
Sluys R 1990 The meaning and implications of genealogical tree diagrams. Z zool Systematik Evolutionsforsch 22:1-8
Sluys R 1991 Species concepts, process analysis, and the hierarchy of nature. Experientia 47: 1162-1170.
Sluys R 1992 A synopsis of Antarctic plagiostomids (Platyhelminthes : Prolecithophora) with the description of a new species and remarks on taxonomy, phylogeny and biogegraphy. Polar Biol 12(5):507-518
Sluys R 1992 Synopsis of the freshwater triclads of the Carribean. Uitgaven Nat Schap Stud Caraib Gebeid 132
Sluys R 1994 Explanations for biogeographic tracks across the Pacific Ocean - a challenge for paleogeography and historical biogeography. Progress in Physical Geography 18: 42-58.
Sluys R 1995 Platyhelminths as paleogeograhical indicators. Hydrobiologia 305:49-53
Sluys R 1996 Reconsiderations of the species status of some South American planarians (Platyhelminthes: Tricladida: Paludicola). Proc Biol Soc Washington 109: 229-235
Sluys R 1996 The notion of homology in current comparative biology. Journal of Zoological Systematics and Evolutionary Research 34: 145-152.
Sluys R 1997 An old problem in a new perspective: the enigmatic evolutionary relationships of some Australian freshwater planarians (Platyhelmithes, Tricladida, Paludicola). Can J Zool 75:459-471
Sluys R 1998 Land planarians (Platyhelminthes, Tricladida, Terricola) in biodiversity and conservation studies. Pedobiologia 42:490-494
Sluys R 1999 Global diversity of land planarians (Platyhelminthes, Tricladida, Terricola): a new indicator-taxon in biodiversity and conservation studies. Biodiversity and Conversation 8: 1663-1681
Sluys R 2001 Towards a phylogenetic classification of dugesiid genera (Platyhelminthes, Tricladida, Dugesiidae): a morphological perspective.. In: Littlewood DTJ & Bray RA, eds. Interrelationships of the Platyhelminthes. Taylor & Francis, London. pp 57-74
Sluys R 2002 Heen en weer: bereisde platwormen. 35  
Sluys R 2007 Annotations on freshwater planarians (Platyhelminthes Tricladida Dugesiidae) from the Afrotropical Region. Tropical Zoology 20:229-257
Sluys R 2012 A new, sibling species of cave flatworm from Switzerland (Platyhelminthes, Tricladida, Dendrocoelidae). Revue Suisse de Zoologie 119: 181-188.
Sluys R 2013 The unappreciated, fundamentally analytical nature of taxonomy and the implications for the inventory of biodiversity. Biodiversity and Conservation 22: 1095-1105.
Sluys R 2016 Invasion of the flatworms. American Scientist 104: 288-295.
[https://doi.org/10. 1511/2016.122.288
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Sluys R 2017 Invasion der Plattwuermer. Spektrum der Wissenschaft 5.17: 32-38.
Sluys R 2017 La invasion de las planarias. Investigacion y Ciencia Mayo 2017: 77-86.
Sluys R 2022 Biogeography and diversification of the freshwater planarian Crenobia alpina sensu lato (Platyhelminthes, Tricladida): A synthetic review. Zoologischer Anzeiger, Volume 300: 12-26
[doi: doi.org/10.1016/j.jcz.2022.05.010
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Sluys R, Ball IR 1983 On the status and identity of Procerodes warreni (Girard,1850) (Turbellaria, Tricladida, Maricola). Mitt Zool Mus Berlin 59:3-10.
Sluys R, Ball IR 1985 On Bothrioplana semperi from Australia, with a review of the species (Platyhelminthes, Proseriata, Bothrioplanidae). The Beagle, N. Terr Mus of A&S 2(1): 129-137
Sluys R, Ball IR 1988 A synopsis of the marine triclads of Australia and New Zealand (Platyhelminthes: Tricladida: Maricola). Invertebr Taxon 2(7): 915-960
Sluys R, Ball IR 1990 The aquatic triclads (Platyhelminthes, Tricladida) of the Bismarck Archipelago. Steenstrupia 16:13-20
Sluys R, Benazzi M 1992 A new finding of a subterranean dendrocoelid flatworm from Italy (Platyhelminthes, Tricladida, Paludicola). Stygologia 7(4):213-217
Sluys R, Berg M van den 1982 On the specific status of the Cyprus pied wheatear Oenanthe-cypriaca. Ornis Scandinavica 13: 123-128.
Sluys R, Bush LF 1988 On Pentacoelum punctatum, an amphi-Atlantic marine triclad (Platyhelminthes: Tricladida: Maricola). Trans. Am. Microsc. Soc. 107(2):162-170
Sluys R, Cannon LRG 1989 A new marine triclad from the west Pacific (Platyhelminthes: Tricladida: Maricola). Invertebrate Taxonomy 3(2):149-154
Sluys R, De Jong H 1984 Chromosome morphological studies of Dugesia gonocephala S.L. (Platyhelminthes, Tricladida). Caryologia 37:9-20
Sluys R, De Vries EJ 1988 The aquatic triclads of the Crozet Islands (Platyhelminthes: Tricladida). [Obrimoposthia, Synsiphonium, Dugesia] Zool J Linn Soc 94: 203-217.
Sluys R, Faubel A, Rajagopal S, van der Velde G 2005 A new and alien species of 'oyster leech' (Platyhelminthes, Polycladida, Stylochidae) from the brackish North Sea Canal, The Netherlands. Helgoland Marine Research 59(4): 310-314
Sluys R, Grant LJ, Blair D 2007 Freshwater planarians from artesian springs in Queensland, Australia (Platyhelminthes, Tricladida, Paludicola). Contributions to Zoology 76(1): 9-19
Sluys R, Hauser J, Wirth QJ 1997 Deviation from the groundplan: a unique new species of freshwater planarian from south Brazil (Platyhelminthes, Tricladida, Paludicola). Journal of Zoology (London) 241(3):593-601
[doi: 10.1111/j.1469-7998.1997.tb04851.x
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Sluys R, Hazevoet CJ 1999 Pluralism in species concepts: dividing nature at its diverse joints. Species Diversity 4: 243-256
Sluys R, Joffe B, Cannon LRG 1995 An alien flatworm in Australian waters. Memoirs of the Queensland Museum 38: 642
Sluys R, Kawakatsu M 1995 Taxon names of the Tricladida Maricola (Platyhelminthes): additions and emendations. Bull Zool Mus Univ Amsterdam 14: 153-160
Sluys R, Kawakatsu M 2000 New southernmost record of Paucumara trigonocephala (Platyhelminthes, Tricladida, Maricola) from Queensland with documentation of earlier records from Japan. Queensland Naturalist 38: 24-30
Sluys R, Kawakatsu M 2001 Contribution to an inventory of the freshwater planarians of Australia and New Zealand (Platyhelminthes, Tricladida, Dugesiidae), with distribution maps of the species examined. Beaufortia / Bull Zool Mus Univ Amsterdam 51(10):163-198
Sluys R, Kawakatsu M 2005 Biodiversity of marine planarians revisited (Platyhelminthes, Tricladida, Maricola). J Nat Hist 39(6):445-467
Sluys R, Kawakatsu M 2005 A replacement name for Ichthyophaga Syromiatnikova, 1949 (Platyhelminthes: Prolecithophora), with a nomenclatural analysis of its avian senior homonym. Species Diversity, Tokyo, 10 (1): 63-68
Sluys R, Kawakatsu M 2006 Towards a phylogenetic classification of dendrocoelid freshwater planarians (Platyhelminthes): a morphological and eclectic approach. Journal of Zoological Systematics and Evolutionary Research 44(4): 274-284
Sluys R, Kawakatsu M, & Ponce de León R 2005 Morphological stasis in an old and widespread group of species: Contribution to the taxonomy and biogeography of the genus Girardia (Platyhelminthes, Tricladida, Paludicola) Studies on Neotropical Fauna and Environment, 40(2):155-180
Sluys R, Kawakatsu M, Bleeker J 2006 The Kawakatsu Collection incorporated within the collections of the Zoological Museum Amsterdam. [Poster presented at 10th International Symposium on Flatworm Biology, Innsbruck, Austria, August, 2006]
Sluys R, Kawakatsu M, Riutort M, Baguñà J 2009 A new higher classification of planarian flatworms (Platyhelminthes, Tricladida) Journal of Natural History 43(29):1763-1777
[doi: doi.org/10.1080/00222930902741669
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Sluys R, Kawakatsu M, Timoshkin OA 2001 Taxonomic redescription of Phagocata sibirica and comparison with Phagocata vivida (Tricladida, Paludicola). Belg J Zool 131(Suppl 1):193-199
Sluys R, Kawakatsu M, Winsor L 1998 The genus Dugesia in Australia, with its phylogenetic analysis and historical biogeography (Platyhelminthes, Tricladida, Dugesiidae). Zool Scr 27: 273-289
Sluys R, Kawakatsu M, Winsor L 1998 The genus Dugesia in Australia, with its phylogenetic analysis and historical biogeoraphy (Platyhelminthes, Tricladida, Dugesiidae). Zool Scripta 27: 273-289
Sluys R, Kawakatsu M, Yamamoto K 2010 Exotic freshwater planarians currently known from Japan. Belgian Journal of Zoology 140: 103-U38
Sluys R, Laumer C 2019 A new genus and species of a terrestrial cavernicolan planarian from Barro Colorado Island, Panama (Platyhelminthes, Tricladida, Cavernicola). Zootaxa, Volume:4586(1): 187-193
Sluys R, Martens K, Schram FR 2004 The Phylocode: naming of biodiversity at a crossroads. Trends Ecol Evol 19:280-281
Sluys R, Mateos E, Riutort M, Alvarez-Presas M 2016 Towards a comprehensive, integrative analysis of the diversity of European microplaninid land flatworms (Platyhelminthes, Tricladida, Microplaninae), with the description of two peculiar new species. Systematics and Biodiversity 14: 9-31.
[doi: doi.org/10.1080/14772000.2015.1103323
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Sluys R, Neumann M, De Lima RF, Drewes RC 2017 Land flatworms (Platyhelminthes, Geoplanidae) of Sao Tome: a first account on their diversity, with the description of five new species. Zootaxa Volume:4221, Issue:3: 291-322
Sluys R, Ribas M, Baguna J 1995 Taxonomy, ecology, and karyology of a new species of Phagocata from Spain, with a discussion on the phylogenetic systematics of the genus Phagocata s.l. (Playhelminthes, Tricladida, Paludicola). Canadian Journal of Zoology - Revue Canadienne de Zoologie 73: 557-568.
[doi: 10.1139/z95-064
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Sluys R, Riutort M 2018 Planarian diversity and phylogeny. Methods in Molecular Biology 1774: 1-56: Humana Press, Springer Science+Business Media.
Sluys R, Riutort M, Ribas M, Baguna J 1990 On Pentacoelum hispaniensis, a marine triclad from Spanish freshwater (Platyhelminthes, Tricladida, Maricola). Hydrobiologia 199:101-106
Sluys R, Rohde K 1991 A new species of freshwater triclad (Platyhelminthes: Tricladida) from Australia. Zool J Linn Soc 102(2):153-162
Sluys R, Schalk PH, Schram FR 1995 Systematic Cirripedology in the 21st century: multimedia biodiversity information systems. 21st century: multimedia biodiversity information systems. 43-48
Sluys R, Smolders I, Kawakatsu M, Pietsch TW, Kuranishi RB 2009 Freshwater Planarians (Platyhelminthes: Tricladida: Planariidae) from the Kuril Islands and Kamchatka Species Diversity 14: 307-322
Sluys R, Solà E, Gritzalis K, Vila-Farré M, Mateos E, Riutort M 2013 Integrative delineation of species of Mediterranean freshwater planarians (Platyhelminthes: Tricladida: Dugesiidae). Zoological Journal of the Linnean Society 169(3): 523–547.
[doi: 10.1111/zoj.12077
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Sluys R, Stocchino GA 2017 Bipolarity in planarians is not induced by space travel. Regeneration (Off) (4): 153-155.
Sluys R, Timoshkin OA, Kawakatsu M 1996 A new species of giant planarian from Lake Baikal, with some remarks on character states in the Dendrocoelidae (Platyhelminthes, Tricladida, Paludicola). 8th Int. Sympos. Biol. Turbellaria
Sluys R, Timoshkin OA, Kawakatsu M 1998 A new species of giant planarian from Lake Baikal, with some remarks on character states in the Dendrocoelidae (Platyhelminthes, Tricladida, Paludicola). Hydrobiologia 383: 69-75 [revised version published in 2006]
Sluys R, van Ginkel W 1989 New records of Fecampia abyssicola (Platyhelminthes: Rhabdocoela: Fecampiidae). Cah Biol Mar 30: 235-241
Sluys R, Vila-Farre M, Alvarez-Presas M, Riutori M, Kawakatsu M, Tulp AS 2015 The diet and distribution of Pentacoelum kazukolinda (Kawakatsu & Mitchell, 1984), a maricolan planarian with a freshwater ecology. Zoologica Scripta, Volume:44(1): 72-91
Sluys R, Vila-Farre M, Rink J, John EJ 2018 An intriguing, new planarian species from Tasmania, with a discussion on protandry in triclad flatworms (Platyhelminthes, Tricladida). Acta Zoologica (Copenhagen), Vol 99(4): 404-414
Solà E, Sluys R, Gritzalis K, Riutort M 2013 Fluvial basin history in the northeastern Mediterranean region underlies dispersal and speciation patterns in the genus Dugesia (Platyhelminthes, Tricladida, Dugesiidae) Molecular Phylogenetics and Evolution, 66(3):877-88
Solà E, Sluys R, Riutort M, Kawakatsu M 2023 Molecular phylogenetics facilitates the first historical biogeographic analysis of the hammerhead worms (Platyhelminthes: Tricladida: Bipaliinae), with the description of twelve new species and two new genera Zootaxa 5335(1)
[doi: 10.11646/zootaxa.5335.1.1
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Solà E, Sluys R, Segev O, Blaustein L, Riutort M 2015 The taxonomic status of Dugesia biblica from Israel and Turkey (Platyhelminthes, Tricladida, Dugesiidae). Zookeys. 2015; (506): 1–12.
Song X-Y, Li W-X, Sluys R, Huang S-X, Li S-F, Wang A-T 2020 A new species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from China, with an account on the histochemical structure of its major nervous system. Zoosystematics and Evolution, Volume 96(2): 431-447
Stocchino GA, Dols-Serrate D, Sluys R, Riutort M, Onnis C, Manconi R 2021 Amphibioplanidae: a new branch and family on the phylogenetic tree of the triclad flatworms (Platyhelminthes: Tricladida), represented by a species from Sardinian caves with a remarkable lifestyle. Zoological Journal of the Linnean Society, zlaa183
[https://doi.org/10.1093/zoolinnean/zlaa183
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Stocchino GA, Manconi R, Corso G, Sluys R, Casu S, Pala M 2009 African planarians: morphology and karyology of Dugesia maghrebiana sp. n. (Platyhelminthes, Tricladida) from Tunisia. Italian Journal of Zoology (Modena) Volume: 76 Issue: 1: 83-91
[doi: doi.org/10.1080/11250000802141683
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Stocchino GA, Sluys R, Montanari A, Manconi R 2017 A new species of stygobiont freshwater planarian (Platyhelminthes, Tricladida) from a chemoautotrophic ecosystem: the Frasassi karst in Italy. Zootaxa Volume:4323 Issue:4 Pages:547-560
Stocchino GA, Sluys R, Deri P, Manconi R 2013 Integrative taxonomy of a new species of planarian from the Lake Ohrid basin, including an analysis of biogeographical patterns in freshwater triclads from the Ohrid region (Platyhelminthes, Tricladida, Dugesiidae). ZooKeys, 313: 25-43
Stocchino GA, Sluys R, Harrath AH, Mansour L, Manconi R 2019 The invasive alien freshwater flatworm Girardia tigrina (Girard, 1850) (Platyhelminthes, Tricladida) in Western Europe: new insights into its morphology, karyology and reproductive biology. Contributions to Zoology, Volume:88(2): 236-256
Stocchino GA, Sluys R, Kawakatsu M, Sarbu SM, Manconi R 2017 A new species of freshwater flatworm (Platyhelminthes, Tricladida, Dendrocoelidae) inhabiting a chemoautotrophic groundwater ecosystem in Romania. European Journal of Taxonomy Volume:342 Pages:1-21
[doi: DOI: https://doi.org/10.5852/ejt.2017.342
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Stocchino GA, Sluys R, Manconi R 2012 A new species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from the Afromontane forest in South Africa, with an overview of freshwater planarians from the African continent Zootaxa 3551: 43-58
Stocchino GA, Sluys R, Manconi R 2014 A new and aberrant species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from Madagascar. ZooKeys, 425: 71-86
Stocchino GA, Sluys R, Marcia P, Manconi R 2013 Subterranean aquatic planarians of Sardinia, with a discussion on the penial flagellum and the bursal canal sphincter in the genus Dendrocoelum (Platyhelminthes, Tricladida, Dendrocoelidae). Journal of Cave and Karst Studies 75: 93–112.
Stocchino GA, Sluys R, Riutort M, Sola E, Manconi R 2017 Freshwater planarian diversity (Platyhelminthes: Tricladida: Dugesiidae) in Madagascar: new species, cryptic species, with a redefinition of character states. Zoological Journal of the Linnean Society, Vol:181 Issue:4:727-756
Tekaya S, Sluys R, Zghal F 1996 Cerbussowia cerruti Wilhelmi, 1909, from the gulf of Tunis (Platyhelminthe, Triclade marin). Archives de l'Institut Pasteur de Tunis 73(3-4): 211-214
Tekaya S, Sluys R, Zghal F 1997 Sperm transfer and fertilization in the marine planarian Sabussowia dioica (Platyhelminthes, Tricladida, Maricola). International Journal of Invertebrate Reproduction and Development 32: 143-147.
Tekaya S, Sluys R, Zghal F 1999 Cocoon production, deposition, hatching, and embryonic development in the marine planarian Sabussowia dioica (Platyhelminthes, Tricladida, Maricola). Invertebrate Reproduction and Development 35: 215-223.
Thielicke M, Sluys R 2019 Prey capture and feeding behaviour in an endemic land flatworm from Sao Tome Island. Journal of Natural History, Vol 53(23-24): 1385-1393
Vasquez-Doorman C, Brusa F, Reyes J, Sluys R, Vila-Farre M, Allende ML 2022 First finds in North and South America of Pentacoelum kazukolinda (Platyhelminthes: Tricladida), a worldwide invasive flatworm. Bioinvasions Records, Vol 11(4): 1078-1094
[https://doi.org/10.3391/bir. 2022.11.4.25
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Vila-Farré M, Mateos E, Sluys R 2006 Preliminary data on the status of triclad flatworms in Spain. Berichte des Naturwissenschaftlichen-Medizinischen Verein Innsbruck Suppl. 16: 96.
Vila-Farre M, Mateos E, Sluys R, Romero R 2008 Terrestrial planarians (Platyhelminthes, Tricladida, Terricola) from the Iberian Peninsula: new records and description of three new species Zootaxa 1739(1): 1-20
[https://doi.org/10.11646/zootaxa.1739.1.1
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Vila-Farré M, Romero R, Sluys R, Fernández J, Sancho-Prat M, Ferrer X 2008 Ecology, karyology, and anatomy of the planarian Pentacoelum hispaniense Sluys, 1989 (Platyhelminthes, Tricladida). Graellsia 64:45-54
Vila-Farré M, Sluys R, Almagro I, Handberg-Thorsager M, Romero R 2011 Freshwater planarians (Platyhelminthes, Tricladida) from the Iberian Peninsula and Greece: diversity and notes on ecology Zootaxa 2779: 1-57
Vila-Farre M, Sluys R, D'Aniello S, Cebria F, Ferrer X, Romero R 2009 Marine planarians (Platyhelminthes: Tricladida: Maricola) from the western Mediterranean Sea and the Cantabrian coast: new records, one new genus, and immunocytochemistry of the nervous system. Journal of the Marine Biological Association of the United Kingdom, pages 1-14
Vila-Farre M, Sluys R, D'Aniello S, Cebria F, Ferrer X, Romero R 2010 Marine planarians (Platyhelminthes: Tricladida: Maricola) from the western Mediterranean Sea and the Cantabrian coast: new records, one new genus, and immunocytochemistry of the nervous system Journal of the Marine Biological Association of the United Kingdom 90: 409-422.
Vila-Farré M, Sluys R, Mateos E, Jones HD, Romero R 2011 Land planarians (Platyhelminthes: Tricladida: Geoplanidae) from the Iberian Peninsula: new records and description of two new species, with a discussion on ecology Journal of Natural History 45(15-16): 869-891
[doi: doi.org/10.1080/00222933.2010.536267
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Wang L, Chen J-z, Dong Z-M, Chen G-W, Sluys R, Liu D-z 2021 Integrative descriptions of two new species of Dugesia from Hainan Island, China (Platyhelminthes, Tricladida, Dugesiidae). ZooKeys, Vol 1059: 89-116
[https://doi.org/10.3897/zookeys.1059.65633
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Wang L, Wang Y, Dong Z, Chen G, Sluys R, Liu D 2022 Integrative taxonomy unveils a new species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from the southern portion of the Taihang Mountains in northern China, with the description of its complete mitogenome and an exploratory analysis of mitoch.. Integrative Biology, Vol 17(6): 1193-1214
[doi: 10.1111/1749-4877.12605
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Wang L, Dong Z-M, Chen G-W, Sluys R, Liu D 2021 Integrative descriptions of two new species of Dugesia from Hainan Island, China (Platyhelminthes, Tricladida, Dugesiidae). ZooKeys 1028: 1-28
[doi: 10.3897/zookeys.1028.60838
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Wang L, Zhu S, Ma F, Li X, Zhao Y, Sun X, Li N, Sluys R, Liu D, Dong Z, Chen G 2024 Two new species of freshwater planarian (Platyhelminthes, Tricladida, Dugesiidae, Dugesia) from southern China exhibit unusual karyotypes, with a discussion on reproduction in aneuploid species Journal of Zoological Systematics and Evolutionary Research 2024: 1-18
[doi: 10.1155/2024/8299436
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Winsor L, Sluys R 2018 Basic histological techniques for planarians. In: Rink, Jochen C., (ed.) Planarian Regeneration: methods and protocols. Methods in Molecular Biology, 1774 . Humana Press, New York, NY, USA, pp. 285-351.
[doi: 10.1007/978-1-4939-7802-1
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Wu S-K, Kawakatsu M, Lue K-Y, Lee J-D, Tsai C-L, Lin H-H, Sluys R, Sasaki G 2005 A preliminary study on land planarians of Taiwan. Endemic Species Research (Taiwain) 7: 23-40.
Yang H-M, Sluys R, Kawakatsu M, Min G-S 2018 New molecular sequences for two genera of marine planarians facilitate determination of their position in the phylogenetic tree, with new records for two species (Platyhelminthes, Tricladida, Maricola). ZooKeys Vol:781: 1-17
Yang Y, Li J-U, Sluys R, Li W-X, Li S-F, Wang A-T 2020 Unique mating behavior, and reproductive biology of a simultaneous hermaphroditic marine flatworm (Platyhelminthes, Tricladida, Maricola). Invertebrate Biology, Volume:139(1): 12282
Zeng Z, Wang J, Sluys R, Guo Z, Sun T, Huang X, Li S, Wang A 2024 Integrative description of a new species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from Shennongjia, Central China Zootaxa 5406(4): 535-550
[doi: 10.11646/zootaxa.5406.4.3
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Zhu Y, Huang J, Sluys R, Liu Y, Sun T, Wang A, Zhang Y 2024 Integrative description of a new species of Dugesia (Platyhelminthes, Tricladida, Dugesiidae) from southern China, with its complete mitogenome and a biogeographic evaluation Zoosystematics and Evolution 100(1): 167-182
[doi: 10.3897/zse.100.114016
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